The 5 S's explained: Solid,
Silver, Smoke, Shaded, Shell
Agouti + Wide Band = Golden series
Agouti + Wide Band + dominant Silver = Silver series
In goldens, wide band refers to the yellow banding of the hair shaft.
Non-agouti + Wide Band = Solid Colour
Non-agouti + Wide Band + dominant Silver = Smoke Pattern
The presence or absence of the Inhibitor (silver) gene does not affect the Wide Band effect - Golden Shaded lack the Inhibitor gene, but have a shading pattern comparable to Silver Shaded cats.
The shaded masks homozygous mackerel tabby as is true with most homozygous shaded ASH, in my experience. The recessive inversion of U_ will be called u and uu allows the cats to express a stripped mackerel (Tm_) or classic tabby pattern . Because both parents are homozygous (shaded - UUTmTm, classic uutbtb) all F1 kittens are unstriped tabbies carrying both mackerel and classic tabby patterns (UuTtb).
If the gene behind shaded ASH's is a single dominant gene U_, it is on a separate allele that "masks" the underlying tabby pattern. Table 2 illustrates the expected inheritance if a single dominant gene, U_ produced unstriped tabbies. For the sake of illustration, a shaded "homozygous" shaded female is bred to a classic male without a shaded background.
The U_ gene is not a true complete dominant, since the heterozygote allows the striping on the head and extremities to be expressed. Partial, incomplete, or co-dominance is probably closer to the truth.
The agouti gene probably works similarly in cats and brown tabbies appear to produce both eumelanin and phaeomelanin similar to the wild type agouti mouse. However, agouti mice are stripeless and mouse genetics are unable to explain how stripes occur in cats. Stripes are remarkable because they represent variable expression of the agouti gene. Both the light and the dark striped areas have multiply banded hair, but the banding frequency varies in these sites. In a classic tabby, in the black stripes areas the hair follicles are coordinated to produce hairs with a black (eumelanin) tip followed by a brown (pheomelanin) band in the undercoat in the case of a brown tabby or no melanin in the case of a silver tabby. In the light areas, the hair follicles collaborate to form a different banding pattern with shorter pigmented tips and higher banding frequency. Here, the hairs are multiply banded, usually black, light, black followed by a light undercoat. As one moves from the back to the belly, the frequency and length of the colored bands decreases.
Shaded Silvers have a mix of three patterns: a single broad band and wide undercoat; a few broad bands and wide undercoat or multiple thin bands (as seen in the background colour of Silver Tabbies).
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